Mechanisms regulating steroid hormone receptor gene expression in the ovine uterus : effects of trophoblast interferon tau /

This research was conducted to determine the cellular and

Bibliographic Details
Main Author: Spencer, Thomas Edward, 1968-
Format: Thesis Book
Language:English
Published: [Place of publication not identified] : [publisher not identified] ; 1995.
Subjects:
Online Access:Link to OAKTrust copy
http://proxy.library.tamu.edu/login?url=http://proquest.umi.com/pqdweb?did=742746241&sid=1&Fmt=2&clientId=2945&RQT=309&VName=PQD
Description
Summary:This research was conducted to determine the cellular and
molecular mechanism(s) whereby ovine interferon tau (IFNR)
regulates steroid hormone receptor gene expression in the
endometrium. Results from the first study indicated that
uterine estrogen receptor (ER) and progesterone receptor (PR)
gene expression is regulated in a tissue and cell-type
specific manner during the estrous cycle and early pregnancy.
In both cyclic and pregnant ewes, PR gene expression
decreased to low levels in all endometrial cell types and was
undetectable in the luminal and superficial glandular
epithelium during late diestrus. In pregnant ewes, the
conceptus blocked increases in ER gene expression that
occurred in the endometrial epithelium of cyclic ewes
undergoing luteolysis. Results from the second study
indicated that intrauterine injections of IFNR into cyclic
ewes suppressed increases in endometrial ER MRNA, ER protein,
OTR density and oxytocin-induced prostaglandin F. (PGF.)
release which occurred in control ewes undergoing luteolysis.
Results from the third study indicated that IFNR does not
stabilize or prevent autologous down-regulation of PR MRNA or
protein expression in the endometrial epithelium of
ovariectomized ewes continously exposed to progesterone.
However, IFNR does suppress increases in endometrial ER MRNA
and protein expression which occurred in control ewes
receiving progesterone alone or progesterone plus estrogen.
Results from the fourth study indicated that IFNR prevented
estrogen-induced luteolysis in cyclic ewes by suppressing
increases in ER MRNA expression in the endometrial
epithelium. Results from the fifth study indicated that
progesterone affects estrogen-induced increases in ER, PR and
OTR expression in the PR-positive deep glandular epithelium
and stroma, whereas IFNT regulates estrogen-induced increases
in expression of those hormone receptors in the PR-negative
luminal and superficial glandular epithelium. Finally,
results of the sixth study indicated that IFNT suppresses
increases in transcription of the endometrial ER and OTR
genes, but has no effect on transcription of the PR or 28S
RRNA genes. Collectively, these studies demonstrate that the
antiluteolytic action of IFNT is to suppress ER gene
expression through a negative-acting transcriptional
mechanism to prevent estrogen-induced increases in OTR gene
transcription. This sequence of events, initiated by IFNT
from the conceptus, would prevent OTR formation in the
endometrial epithelium, production of luteolytic pulses of
PGF, and ensure maintenance of the corpus luteum and
successful maternal recognition of pregnancy.
Item Description:Vita.
"Major Subject: Physiology of Reproduction".
Physical Description:xiii, 271 leaves : illustrations ; 28 cm.
Issued also on microfiche from University Microfilms Inc.
Bibliography:Includes bibliographical references.